Based on my own and others' research
SystemID NES
0051 b u 0 RAM Game Mode
0049 b u 0 RAM Frame Counter
0069 b u 0 RAM Hero Action
00D4 b h 0 RAM Screen Scroll 2
00E5 b h 0 RAM Screen Scroll 1
04A5 b u 0 RAM Boss HP
04A6 b u 0 RAM Hero HP
0058 b u 0 RAM Current Stage
0531 b h 0 RAM Score 5
0532 b h 0 RAM Score 4
0533 b h 0 RAM Score 3
0534 b h 0 RAM Score 2
0535 b h 0 RAM Score 1
0021 b u 0 RAM Hero Action Timer
0094 b u 0 RAM Hero Screen Pos X
00B6 b u 0 RAM Hero Pos Y
036A b u 0 RAM Hero Air Mode
0008 b u 0 RAM Game Submode
0378 b u 0 RAM Shrug Counter
002B b u 0 RAM Enemy 1 Action Timer
002C b u 0 RAM Enemy 2 Action Timer
002D b u 0 RAM Enemy 3 Action Timer
002E b u 0 RAM Enemy 4 Action Timer
008E b u 0 RAM Enemy 1 Pos X
008F b u 0 RAM Enemy 2 Pos X
0090 b u 0 RAM Enemy 3 Pos X
0091 b u 0 RAM Enemy 4 Pos X
0080 b h 0 RAM Enemy 1 Action
0081 b h 0 RAM Enemy 2 Action
0082 b h 0 RAM Enemy 3 Action
0083 b h 0 RAM Enemy 4 Action
0374 b u 0 RAM Grab Counter
0090 b h 0 RAM Boss Pos X
004E b u 0 RAM Boss Action
0093 b u 0 RAM Boss Pos X